John Kappler

John Wayne Kappler (born December 22, 1943, in Baltimore, Maryland) is a professor in the Department of Integrated Immunology at National Jewish Health. His principal research is in T cell biology, a subject he collaborates on with his wife Philippa Marrack. In 1983 they discovered the T cell receptor, together with Ellis Reinherz and James Allison.

Awards 1986 – Appointed investigator, Howard Hughes Medical Institute 1989 – Elected member, National Academy of Sciences 1993 – Cancer Research Institute William B. Coley Award 1993 - Paul Ehrlich and Ludwig Darmstaedter Prize 1994 – Louisa Gross Horwitz Prize for Biology or Biochemistry (Columbia University) 2015 – Wolf Prize in Medicine