Charlotte M. Taylor

Dr. Charlotte M. Taylor is a botanist and professor specialising in taxonomy and conservation. She works with the large plant family Rubiaceae, particularly found in the American tropics and in the tribes Palicoureeae and Psychotrieae. This plant family is an economically important group, as it includes plant species used to make coffee and quinine. Taylor also conducts work related to the floristics of Rubiaceae and morphological radiations of the group. Taylor has collected plant samples from many countries across the globe, including Chile, Colombia, Costa Rica, Panama, and the United States of America, She has 172 publications on Research Gate alone.

Education Taylor holds a B.S. from the University of Michigan (1978), and an M.S. (1982) and Ph.D. (1987) from Duke University. In addition to the numerous plants that she has named, Taylor has also conducted taxonomic work and transferred species names between different genera. For this reason, she is linked as an author to 1,091 plant species name citation records through the International Plant Names Index (IPNI). A full list of all 1,091 records can be viewed here.

Rubiaceae projects In addition to her taxonomic work with this family, Taylor maintains two websites related Rubiaceae to the Missouri Botanical Garden website. The Selected Rubiaceae Tribes and Genera website includes taxonomic parts of previously published works related to the family. The content of the website is also incorporated in Tropicos, the online database of taxonomic information about plants maintained and populated by the Missouri Botanical Garden and its scientific staff.

Personal life Taylor attributes her interest in plants to her parents, who were "serious bird watchers." However, she opted to study plants instead of birds because it afforded her more freedom to keep to her own research schedule. an Assistant Curator at the Missouri Botanical Garden. Gereau's research interests include plant nomenclature, floristics and phytogeography of eastern Africa, plant conservation assessment in eastern Africa and in Africa generally, classification and identification of East African flowering plant genera, and taxonomy and systematics of African Sapindaceae. Taylor and Gereau have published together on botanical topics.

New species described As of 2015, Taylor had named 278 plant taxa new to science

Alibertia premontana C.M. Taylor Alseis costaricensis C.M. Taylor Amphidasya amethystina J.L. Clark & C.M. Taylor Amphidasya brevidentata C.M. Taylor Amphidasya elegans C.M. Taylor Amphidasya panamensis C.M. Taylor Appunia megalantha C.M. Taylor & Lorence Bouvardia costaricensis C.M. Taylor Carapichea verrucosa C.M. Taylor Chiococca caputensis Lorence & C.M. Taylor Chomelia chiquitensis C.M. Taylor Chomelia costaricensis C.M. Taylor Chomelia rubra Lorence & C.M. Taylor Chomelia torrana C.M. Taylor Chomelia venulosa W.C. Burger & C.M. Taylor Coussarea acrensis C.M. Taylor Coussarea amplifolia C.M. Taylor Coussarea antioquiana C.M. Taylor Coussarea boliviensis C.M. Taylor Coussarea brevipedunculata C.M. Taylor Coussarea camposiana C.M. Taylor Coussarea cephaeloides C.M. Taylor Coussarea cephaëloides C.M. Taylor Coussarea dulcifolia D.A. Neill, Cerón & C.M. Taylor Coussarea duplex C.M. Taylor Coussarea ecuadorensis C.M. Taylor Coussarea grandifructa C.M. Taylor Coussarea izabalensis C.M. Taylor Coussarea linearis C.M. Taylor Coussarea loftonii subsp. calimana C.M. Taylor Coussarea loftonii subsp. occidentalis C.M. Taylor Coussarea maranonensis C.M. Taylor Coussarea mexiae C.M. Taylor Coussarea nigrescens C.M. Taylor & Hammel Coussarea pilosula C.M. Taylor Coussarea pilosula C.M. Taylor Coussarea pseudopilosula C.M. Taylor Coussarea psychotrioides C.M. Taylor & Hammel Coussarea resinosa C.M. Taylor Coussarea sancti-ciprianii C.M. Taylor Coussarea spiciformis C.M. Taylor Coussarea spiciformis C.M. Taylor Coussarea vasqueziana C.M. Taylor Coutarea coutaportloides C.M. Taylor Coutarea fuchsioides C.M. Taylor Danais antilahimenae C.M. Taylor Danais disticha C.M. Taylor Danais laciniata C.M. Taylor Danais masoalana C.M. Taylor Danais rakotovaoi C.M. Taylor Danais randrianaivoi C.M. Taylor Duroia laevis Devia, C.H. Perss. & C.M. Taylor Duroia sancti-ciprianii Devia, C.H. Perss. & C.M. Taylor Elaeagia chiriquina C.M. Taylor Elaeagia glossostipula C.M. Taylor Faramea accumulans C.M. Taylor Faramea ampla C.M. Taylor Faramea angusta C.M. Taylor Faramea areolata C.M. Taylor Faramea calimana C.M. Taylor Faramea calyptrata C.M. Taylor Faramea coffeoides C.M. Taylor Faramea colombiana C.M. Taylor Faramea condorica C.M. Taylor Faramea correae C.M. Taylor Faramea cupheoides C.M. Taylor Faramea frondosa C.M. Taylor Faramea guaramacalensis C.M. Taylor Faramea longistipula C.M. Taylor Faramea melicoccoides C.M. Taylor Faramea monsalveae C.M. Taylor Faramea ortiziana C.M. Taylor Faramea permagnifolia Dwyer ex C.M. Taylor Faramea robusta C.M. Taylor Faramea sanblasensis C.M. Taylor Faramea schunkeana C.M. Taylor Faramea uncinata C.M. Taylor Faramea vasquezii C.M. Taylor Faramea verticillata C.M. Taylor Gaertnera arenarioides C.M. Taylor Gaertnera breviflora C.M. Taylor Gaertnera hirsuta C.M. Taylor Gaertnera laevis C.M. Taylor Gaertnera littoralis C.M. Taylor Gaertnera malcomberiana C.M. Taylor Gaertnera masoalana C.M. Taylor Gaertnera nitida C.M. Taylor Gaertnera rakotovaoana C.M. Taylor Gaertnera razakamalalana C.M. Taylor Gaertnera robusta C.M. Taylor Gaertnera rubra C.M. Taylor Gaertnera sclerophylla C.M. Taylor Gaertnera velutina C.M. Taylor Gaertnera vernicosa C.M. Taylor Gaertnera xerophila C.M. Taylor Gonzalagunia mildrediae D.R. Simpson ex C.M. Taylor Gonzalagunia osaensis C.M. Taylor Gonzalagunia villosa D.R. Simpson ex C.M. Taylor Guettarda subcapitata C.M. Taylor Hillia grayumii C.M. Taylor Hillia oaxacana C.M. Taylor Hillia pumila C.M. Taylor Hillia rivalis C.M. Taylor Hillia subg. Andinae C.M. Taylor Hillia subg. Illustres C.M. Taylor Hillia subg. Tetrandrae C.M. Taylor Hippotis stellata C.M. Taylor & Rova Hoffmannia barbillana C.M. Taylor Hoffmannia boliviana C.M. Taylor Hoffmannia boraginoides Dwyer ex W.C. Burger & C.M. Taylor Hoffmannia coriacea C.M. Taylor Hoffmannia costaricensis C.M. Taylor Hoffmannia formicaria C.M. Taylor Hoffmannia hammelii C.M. Taylor Hoffmannia limonensis C.M. Taylor Hoffmannia micrantha C.M. Taylor Hoffmannia pacifica C.M. Taylor Hoffmannia pseudovesiculifera C.M. Taylor Hoffmannia rivalis C.M. Taylor Hoffmannia subcapitata C.M. Taylor Hoffmannia tilaranensis C.M. Taylor Hoffmannia turrialbana C.M. Taylor Hoffmannia veraguensis C.M. Taylor Ixora knappiae C.M. Taylor Joosia antioquiana C.M. Taylor Joosia capitata C.M. Taylor Joosia confusa C.M. Taylor Joosia frondosa C.M. Taylor Ladenbergia franciscana C.M. Taylor Malanea campylocarpa C.M. Taylor Malanea cylindrica C.M. Taylor Malanea ecuadorensis C.M. Taylor Manettia longipedicellata C.M. Taylor Margaritopsis inconspicua C.M. Taylor Monnina ferreyrae C.M. Taylor Monnina parasylvatica C.M. Taylor Neoblakea ecuadorensis C.M. Taylor Notopleura acuta C.M. Taylor Notopleura aequatoriana C.M. Taylor Notopleura albens C.M. Taylor Notopleura amicitiae C.M. Taylor Notopleura bahiensis C.M. Taylor Notopleura biloba C.M. Taylor Notopleura bryophila C.M. Taylor Notopleura callejasii C.M. Taylor Notopleura capitata C.M. Taylor Notopleura cincinalis C.M. Taylor Notopleura cocleensis C.M. Taylor Notopleura congesta C.M. Taylor Notopleura corniculata C.M. Taylor Notopleura corymbosa C.M. Taylor Notopleura costaricensis C.M. Taylor Notopleura cundinamarcana C.M. Taylor Notopleura elegans C.M. Taylor Notopleura episcandens C.M. Taylor & Lorence Notopleura hondurensis C.M. Taylor Notopleura hurtadoi C.M. Taylor Notopleura hypolaevis C.M. Taylor Notopleura iridescens C.M. Taylor Notopleura lanosa C.M. Taylor Notopleura longiflora C.M. Taylor Notopleura longipedunculata C.M. Taylor Notopleura montana C.M. Taylor Notopleura multinervia C.M. Taylor Notopleura nepokroeffiae C.M. Taylor Notopleura obtusa C.M. Taylor Notopleura pacorana C.M. Taylor Notopleura parasiggersiana C.M. Taylor Notopleura parvifolia C.M. Taylor Notopleura penduliflora C.M. Taylor Notopleura pilosula C.M. Taylor Notopleura pyramidata C.M. Taylor Notopleura recondita Hammel & C.M. Taylor Notopleura sanblasensis C.M. Taylor Notopleura scarlatina C.M. Taylor Notopleura spiciformis C.M. Taylor Notopleura steyermarkiana C.M. Taylor Notopleura submarginalis C.M. Taylor Notopleura torrana C.M. Taylor Notopleura triaxillaris C.M. Taylor Notopleura tubulistipula C.M. Taylor Notopleura vargasiana C.M. Taylor Notopleura zarucchiana C.M. Taylor Paederia taolagnarensis Razafim. & C.M. Taylor Palicourea albaniana C.M. Taylor Palicourea albocaerulea C.M. Taylor Palicourea anderssoniana C.M. Taylor Palicourea andina C.M. Taylor Palicourea andina subsp. panamensis C.M. Taylor Palicourea anianguana C.M. Taylor Palicourea asplundii C.M. Taylor Palicourea australis C.M. Taylor Palicourea awa C.M. Taylor Palicourea azulina C.M. Taylor Palicourea azurea C.M. Taylor Palicourea bellula C.M. Taylor Palicourea betancuriana C.M. Taylor Palicourea bullulata C.M. Taylor Palicourea cajamarcana C.M. Taylor Palicourea calophlebioides C.M. Taylor Palicourea canarina C.M. Taylor Palicourea candida C.M. Taylor Palicourea chignul C.M. Taylor Palicourea clerodendroides C.M. Taylor Palicourea cogolloi C.M. Taylor Palicourea condorica C.M. Taylor Palicourea corniculata C.M. Taylor Palicourea cornigera C.M. Taylor Palicourea crystallina C.M. Taylor Palicourea cuspidata subsp. occidentalis C.M. Taylor Palicourea cutucuana C.M. Taylor Palicourea deviae C.M. Taylor Palicourea diminuta C.M. Taylor Palicourea dodsoniana C.M. Taylor Palicourea eburnea C.M. Taylor Palicourea foreroi C.M. Taylor Palicourea frontinoensis Cogollo & C.M. Taylor Palicourea garcioides C.M. Taylor Palicourea gelsemiiflora C.M. Taylor Palicourea gemmiflora C.M. Taylor Palicourea gentryi C.M. Taylor Palicourea glandulifera C.M. Taylor Palicourea gomezii C.M. Taylor Palicourea grandiceps C.M. Taylor Palicourea hammelii C.M. Taylor Palicourea harlingii C.M. Taylor Palicourea hollinensis C.M. Taylor Palicourea hondurensis C.M. Taylor Palicourea ianthina C.M. Taylor Palicourea jaramilloi C.M. Taylor Palicourea jatun-sachensis C.M. Taylor Palicourea kahirica C.M. Taylor Palicourea lemoniana C.M. Taylor Palicourea lewisiorum C.M. Taylor Palicourea locellata C.M. Taylor Palicourea loxensis C.M. Taylor Palicourea lozanoana C.M. Taylor Palicourea lugoana C.M. Taylor Palicourea luteonivea C.M. Taylor Palicourea macarthurorum C.M. Taylor Palicourea madidiensis C.M. Taylor Palicourea matamana C.M. Taylor Palicourea meieri C.M. Taylor Palicourea mistratoana C.M. Taylor Palicourea murciae C.M. Taylor Palicourea neillii C.M. Taylor Palicourea orosiana C.M. Taylor Palicourea orquidea C.M. Taylor Palicourea otongaensis C.M. Taylor Palicourea oxapampana C.M. Taylor Palicourea pachystipula C.M. Taylor Palicourea palaciosii C.M. Taylor Palicourea palustris A.C. Gilman & C.M. Taylor Palicourea parrana C.M. Taylor Palicourea paujilensis C.M. Taylor Palicourea pendula C.M. Taylor Palicourea pereziana C.M. Taylor Palicourea plowmanii D.R. Simpson ex C.M. Taylor Palicourea premontana C.M. Taylor Palicourea prodiga Standl. ex C.M. Taylor Palicourea providenciana J. Sánchez-Gonz. & C.M. Taylor Palicourea purpurea C.M. Taylor Palicourea quadrifolia subsp. leticiana C.M. Taylor Palicourea quadrilateralis C.M. Taylor Palicourea quinquepyrena C.M. Taylor Palicourea rodriguezii C.M. Taylor Palicourea roseofaucis C.M. Taylor Palicourea sancti-ciprianii C.M. Taylor Palicourea sanluisensis C.M. Taylor Palicourea sect. Cephaeloides C.M. Taylor Palicourea sect. Chocoanae C.M. Taylor Palicourea sect. Crocothyrsae C.M. Taylor Palicourea sect. Didymocarpae C.M. Taylor Palicourea sect. Grandiflorae C.M. Taylor Palicourea sect. Montanae C.M. Taylor Palicourea sect. Obovoideae C.M. Taylor Palicourea sect. Pseudoamethystinae C.M. Taylor Palicourea sect. Psychotrioides C.M. Taylor Palicourea shuar C.M. Taylor Palicourea simpsonii C.M. Taylor Palicourea skotakii C.M. Taylor Palicourea smithiana C.M. Taylor Palicourea sopkinii C.M. Taylor Palicourea spathacea C.M. Taylor Palicourea stellata C.M. Taylor Palicourea subalatoides C.M. Taylor Palicourea subg. Montanae C.M. Taylor Palicourea subtomentosa subsp. lojana C.M. Taylor Palicourea sucllii C.M. Taylor Palicourea tilaranensis C.M. Taylor Palicourea topoensis C.M. Taylor Palicourea tunquiensis C.M. Taylor Palicourea ucayalina C.M. Taylor Palicourea ulloana C.M. Taylor Palicourea valenzuelana C.M. Taylor Palicourea vernicosa C.M. Taylor Palicourea vulcanalis Standl. ex C.M. Taylor Palicourea yanesha C.M. Taylor Palicourea zarucchii C.M. Taylor Patima minor C.M. Taylor Pentagonia angustifolia C.M. Taylor Pentagonia australis C.M. Taylor & Janovec Pentagonia dwyeriana C.M. Taylor Pentagonia involucrata C.M. Taylor Pentagonia lobata C.M. Taylor Pentagonia monocaulis C.M. Taylor Pentagonia osaensis C.M. Taylor Pentagonia sanblasensis C.M. Taylor Pittoniotis rotata C.M. Taylor Posoqueria chocoana C.M. Taylor Posoqueria correana C.M. Taylor Posoqueria costaricensis C.M. Taylor Posoqueria grandifructa Hammel & C.M. Taylor Posoqueria laevis C.M. Taylor Posoqueria longifilamentosa C.M. Taylor Posoqueria robusta Hammel & C.M. Taylor Posoqueria tarairensis C.M. Taylor & Cortés-Ballén Psychotria acicularis C.M. Taylor Psychotria anisophylla C.M. Taylor Psychotria areolata C.M. Taylor Psychotria awa C.M. Taylor Psychotria burgeri C.M. Taylor Psychotria calidicola C.M. Taylor Psychotria calyptrata C.M. Taylor Psychotria cauligera C.M. Taylor Psychotria cenepensis C.M. Taylor Psychotria ceronii C.M. Taylor Psychotria cerrocoloradensis Dwyer ex C.M. Taylor Psychotria chamelaensis C.M. Taylor & Domínguez Licona, Eduardo Psychotria chocoana C.M. Taylor Psychotria cochabambana C.M. Taylor Psychotria convergens C.M. Taylor Psychotria cornejoi C.M. Taylor Psychotria cutucuana C.M. Taylor Psychotria davidsmithiana C.M. Taylor Psychotria deneversii C.M. Taylor Psychotria diminuta C.M. Taylor Psychotria esmeraldana C.M. Taylor Psychotria fusiformis C.M. Taylor Psychotria gaitalensis C.M. Taylor Psychotria grahamii C.M. Taylor Psychotria hamiltoniana C.M. Taylor Psychotria hemisphaerica Dwyer ex C.M. Taylor Psychotria herrerana C.M. Taylor Psychotria huampamiensis C.M. Taylor Psychotria hypochlorina C.M. Taylor Psychotria jefensis Dwyer ex C.M. Taylor Psychotria juarezana C.M. Taylor & Lorence Psychotria leitana C.M. Taylor Psychotria lorenciana C.M. Taylor Psychotria matagalpensis C.M. Taylor Psychotria maynasana C.M. Taylor Psychotria monsalveae C.M. Taylor Psychotria montivaga C.M. Taylor Psychotria orosioides C.M. Taylor Psychotria ortiziana C.M. Taylor Psychotria osaensis C.M. Taylor Psychotria ovatistipula C.M. Taylor Psychotria ownbeyi Standl. ex C.M. Taylor Psychotria paeonia C.M. Taylor Psychotria pandoana C.M. Taylor Psychotria paradichroa C.M. Taylor Psychotria paravillosa C.M. Taylor Psychotria purpureocapitata Dwyer ex C.M. Taylor Psychotria puyoana C.M. Taylor Psychotria rhombibractea C.M. Taylor & M.T. Campos Psychotria rhombibracteata C.M. Taylor & M.T. Campos Psychotria romolerouxiana C.M. Taylor Psychotria sacciformis C.M. Taylor Psychotria saltatrix C.M. Taylor Psychotria sanblasensis C.M. Taylor Psychotria sanchezii C.M. Taylor Psychotria simiarum subsp. chiapensis C.M. Taylor Psychotria sinuata C.M. Taylor Psychotria soejartoi C.M. Taylor Psychotria sopkinii C.M. Taylor Psychotria tapantiensis C.M. Taylor Psychotria ucumariana C.M. Taylor Psychotria vesiculifera C.M. Taylor Raritebe axillare C.M. Taylor Remijia uniflora C.M. Taylor Rosenbergiodendron reflexum C.M. Taylor & Lorence Rudgea grandifructa C.M. Taylor & M. Monsalve Rudgea hemisphaerica Dwyer ex C.M. Taylor Rudgea laevis C.M. Taylor Rudgea mandevillifolia Dwyer ex C.M. Taylor Rudgea mcphersonii Dwyer ex C.M. Taylor Rudgea sanblasensis C.M. Taylor Rustia kosnipatana S. Will & C.M. Taylor Sabicea chocoana C.M. Taylor Schizocalyx condoricus D.A. Neill & C.M. Taylor Schizocalyx truncatus C.M. Taylor Schradera obtusifolia C.M. Taylor Schradera reticulata C.M. Taylor ex J. Sánchez-Gonz. Simira cesariana C.M. Taylor Simira hirsuta C.M. Taylor Warszewiczia uxpanapensis subsp. meridionalis C.M. Taylor

Works Selected publications on Palicourea taxonomy Taylor, C.M. 1997. Conspectus of the genus Palicourea (Rubiaceae: Psychotrieae) with the description of same new species from Ecuador. Ann. Missouri Bot. Gard. 84: 224-262. Taylor, C.M, D.H. Lorence, & R.E. Gereau. 2010. Rubiacearum americanarum magna hama pars XXV: The nocturnally flowering Psychotria domingensis-Coussarea hondensis group plus three other Mesoamerican Psychotria species transfer to Palicourea. Novon 20: 481-492.

Selected Publications on Genera of Palicoureeae and Psychotrieae Taylor, C.M. 2001. Overview of the neotropical genus Notopleura (Rubiaceae: Psychotrieae), with the description of some new species. Ann. Missouri Bot. Gard. 88: 478-515. Taylor, C.M. 2003. Rubiacearum americanarum magna hama. Pars XIV. New species and a new combination in Notopleura (Psychotrieae) from Central and South America. Novon 13: 228-260. Taylor, C.M. 2005. Margaritopsis (Rubiaceae, Psychotrieae) in the Neotropics. Syst. Geogr. Pl. 75: 161-177. Taylor, C.M. 2011. The genus Coccochondra (neotropical Rubiaceae) expanded. Pl. Ecol. & Evol. 144(1): 115-118. Taylor, C.M. & R.E. Gereau. 2013. The genus Carapichea (Rubiaceae). Ann. Missouri Bot. Gard. 99: 100-127. Razafimandimbison, S.G., C.M. Taylor, N. Widstrom, T. Paillier, A. Khobadandeh, B. Bremer. 2014. Phylogeny and generic limits in the sister tribes Psychotrieae and Palicoureeae (Rubiaceae): Evolution of schizocarps in Psychotria and origins of bacterial leaf nodules of the Malagasy species. Amer. J. Bot. 101: 1102-1126. Taylor, C.M. 2015. Rubiacearum americanarum magna hama XXXIII: Overview of the new group Palicourea sect. Didymocarpae with four new species and two new subspecies (Palicoureeae). Novon 23: 452-478. Taylor, C.M. 2015. Rubiacearum americanarum magna hama XXXIV: Overview of the new group Palicourea sect. Tricephalium with either new species and a new subspecies (Palicoureeae). Novon 24: 55-95.

Selected flora treatments Taylor, C.M. 1999. Faramea, Coussarea, Palicourea. In: G. Harling and L. Andersson, eds. Flora of Ecuador 62(3): 134-235, 245-314. Taylor, C.M., J.A. Steyermark, P. Delprete, C. Persson, C. Costa, A. Vincentini, & R. Cortés. 2004. Rubiaceae. In: J.A. Steyermark, P.E. Berry, K. Yatskievych, & B.K. Holst, eds. Flora of the Venezuelan Guayana 8:497-847. Campos, M.T.V.A., C.M. & Taylor & D. Zappi. 2007. Rubiaceae. In: M. Hopkins et al., Flora of the Ducke Reserve. Rodriguesia 58: 549-616. Chen, T., H. Zhu; J.-R. Chen; C.M.Taylor, F. Ehrendorfer, H. Lantz, A.M. Funsten, C. Puff. 2011. Rubiaceae. In: Z.-G. Wu, P.H. Raven, & D.-Y. Hong, dirs. Flora of China 19: 57-368. D.H. Lorence, C.M. Taylor, & Collaborators. 2012. Rubiaceae. In: Flora Mesoamericana 4(2): 1-288. Taylor, C.M., B.E. Hammel, & D.H. Lorence. 2014. Rubiaceae. In: B.E. Hammel, M.H. Grayum, M.C. Herrera Mora, & N. Zamora, eds. Manual de Plantas de Costa Rica. Volumen VII. Dicotiledóneas (Picramniaceae-Rutaceae). Monogr. Syst. Bot. Missouri Bot. Gard. 129: 464-779.

Selected floristic catalogues Taylor, C.M. & A. Pool. 1993. Rubiaceae. In: L. Brako et al., Catalogue of the Flowering Plants and Gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45: 1010 1053. Taylor, C.M. 1999. Rubiaceae. In P. Jørgensen & S. León, Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard.75: 855-878. Bacigalupo, N.M., E.L. Cabral, & C.M. Taylor. 2008. Rubiaceae. In: F.O. Zuloaga, O. Morrone, & M.J. Belgrano, eds. Católogo de las Plantas Vasculares del Cono Sur. Monogr. Syst. Bot. Missouri Bot. Gard. 107(3): 1871-2920. Taylor, C.M. 2011. Rubiaceae. In: A. Idárraga P., R. del C. Ortiz, R. Callejas P., & M. Merello. Flora de Antioquia, Catálogo de las Plantas Vasculares, Vol. 2, Listado de las Plantas Vasculares del Departamento de Antioquia, pp. 822–854. Universidad de Antioquia. Taylor, C.M., E. Cabral, & N. Bacigalupo. 2014. Rubiaceae. In: P. Jørgensen, M. Nee, & S. Beck, eds. Catálogo de las Plantas Vasculares de Bolivia. Monogr. Syst. Bot. Missouri Bot. Gard. 127(2): 1140-1171.